Titania's surface is dominated by four major continents, collectively accounting for approximately 54% of the planet’s landmass. The rest of the surface is primarily oceanic. Titania is accompanied by three natural satellites (Taomedon, Themis, Telecleia ), each contributing to tidal activity and likely influencing the planet’s axial stability and climate systems.

The dominant autotrophs on Titania resemble terrestrial vascular plants, although they diverge significantly in taxonomy and evolutionary history. The surface biosphere includes a diverse array of arboreal and floral analogs, but with surprisingly few species of grasses, limited shrub-like organisms, and a near-absence of fungoid decomposers. This imbalance is hypothesized to stem from nutrient monopolization by fast-growing, resource-intensive trees and flowering species, inhibiting the proliferation of other ground-layer vegetation.

Though visually rich, Titania's flora exhibits limited evolutionary complexity, suggesting the planet is in a Mesozoic-equivalent phase of ecological development, akin to Earth's Late Triassic to Jurassic period. Despite their aesthetic appeal, these lifeforms demonstrate little in terms of biochemical or structural innovation when compared to advanced terrestrial flora.

Beneath the surface, Titania's aquatic biomes are dominated by a monoculture of seagrass-like organisms that span vast regions of the ocean floor. While genetically uniform across regions, these aquatic autotrophs display significant morphological plasticity based on local environmental conditions such as temperature, depth, and salinity.

Perhaps the most remarkable marine phenomenon is the prevalence of sessile, flower-like organisms embedded within seagrass meadows. Though reminiscent of Earth corals or sea anemones, these are neither true cnidarians nor plants, and appear to represent a primitive phylum of soft-bodied marine polyps—a convergent evolutionary development unique to Titania.

Terrestrial animal life on Titania is heavily skewed toward arthropod analogs. Insects dominate nearly all ecological niches, from detritivores to apex predators. Due to high atmospheric oxygen levels and relatively mild predatory pressure, many of these organisms have achieved gigantism, paralleling Earth's Carboniferous period in terms of insect scale and diversity.

Other metazoans exist, particularly in aquatic environments, where a limited range of fish-like species have evolved. Most vertebrate analogs are still in nascent stages of development, but notable exceptions have occurred.

Among these exceptions are the Beasten, a lineage of highly adapted endothermic vertebrates that appear to have undergone a rapid evolutionary radiation, possibly due to niche openings or symbiotic relationships. Beasten possess characteristics consistent with mammalian analogs, including high intelligence, social structures, and tool use, making them one of the dominant sentient species on the planet.